Apes
Apes and humans differ from all of the other primates in that they lack external tails. They also are more intelligent and more dependent for survival on learned behavior patterns. The apes and humans are members of the same superfamily, the Hominoidea. Until the last few years, humans were separated into their own family within this superfamily because it was believed that we are significantly different from the apes. However, recent genetic studies and discoveries from the fossil record have made it clear that some of the apes are more similar to humans than previously believed. Subsequently, the living hominoids are now commonly classified into only two families with humans grouped with the great apes:
1. Hylobatidae (gibbons
)
2. Hominidae (orangutans
, gorillas
, chimpanzees
, and bonobos
, and humans)
Gibbons and orangutans live in Southeast Asia, while gorillas, chimpanzees, and bonobos are exclusively African apes. Humans originated in Africa as well.
Natural range of the apes
Asian ApesThe smallest and the most arboreal apes are the 12-13 species of gibbons. Because of their diminutive size, these members of the family Hylobatidae are also referred to as the "lesser apes." Most adult gibbons are only about 3 feet (90 cm.) tall standing upright and 12-20 pounds (5.5-9 kg.) in weight. Males in the biggest gibbon species, known as siamangs
, are up to 30 pounds (13.5 kg.) and have longer arms. Siamangs are different enough from other gibbons to be in their own genus. All gibbons are very slender. Long bushy hair on their bodies makes them look stockier than they actually are. Unlike all of the larger ape species, gibbons have little sexual dimorphism in body size.
The long arms, permanently curved fingers, and light bodies of gibbons make them excellent brachiators
. That is, they move around in trees by swinging under branches with a hand over hand motion. This is also referred to as suspensory climbing. At times, gibbons also walk bipedally, or two footed, on top of branches. However, they are more efficient at brachiation, and 90% of their locomotion is by this means. Each swing can transport a gibbon 20 feet (6 m.) at speeds approaching 35 miles (56 km.) an hour.
Siamang
brachiating
Adult male and female siamang pair
vocalizing in a duet--sound recording
by Dennis O'Neil
(length = 57 secs.)
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Gibbon Gibbons are monogamous
in their mating patterns and form nuclear family groups. That is to say, their communities consist of a single mating pair of adults with their juvenile offspring. They live in well defined territories in the tree tops and rarely go down to the forest floor. Adults regularly defend their territory against others of their species with piercingly loud whooping and hooting vocalizations, much like the indris of Madagascar and the howler monkeys of the New World. Though the calls of the latter two primates sound very different. The calls of different gibbon species are different from each other as well.
Orangutans are the largest and the rarest of the Asian apes. Males often grow to 175-200 pounds (80-90 kg.) and 4½ feet (1.4 m.) tall. At this size, they are usually too large to cross from one tree to another by the branches and must go down to the ground and walk quadrupedally between them. There is marked sexual dimorphism among the orangutans. Males have huge fleshy pads framing the upper part of their faces. In addition, females weigh only about half as much as the males (73-99 lbs or 33-45 kg.). Being lighter, females and juveniles often stay in the trees and use a leaning form of brachiation--they carefully shift their body weight to bend a supporting branch and then grab the next one before the first one breaks.
Adult female orangutan Adult male orangutan Leaning form
of brachiation
(young orangutan)Orangutan mother with
child "fishing out" food
from a rock crevice
Orangutans are intelligent and generally peaceful animals. Most of the time, they live solitary lives browsing fruits and leaves. Some also create simple stick tools to get honey out of bee hives in tree crevices.
Unfortunately, the orangutans are in danger of extinction in the wild because they are hunted for the illegal international pet trade. Their forest territories are also being rapidly cut down for the lumber and cleared by burning for farming, especially in Indonesia where most of them live. According to Conservation International, the 15,000 remaining Indonesian orangutans are currently disappearing at a rate of 1,000 per year. When population size decreases, there is a corresponding decrease in genetic diversity. Species that have little genetic diversity are more easily driven to extinction by a changing environment. The loss of habitat has decimated gibbons in Indonesia as well. However, they are not at as much a risk of becoming extinct because their range extends widely over Southeast Asia.
Kalimantan's Orangutans--video clip from National Geographic Society
(length = 2 mins. 18 secs.)
African Apes
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Male lowland gorilla The largest apes are the gorillas of Africa. Adult males are up to 6 feet (1.8 m.) tall and have 9-10 foot (2.7-3.0 m.) arm spans. They have massive heads with heavy, thick muscles on top that are used to close crushing jaws. Their bodies are stout and very muscular.
Knuckle walking
(subadult male lowland gorilla)Like humans, gorillas are terrestrial animals. They are quadrupedal
knuckle walkers. That is, they walk on the soles of their feet but not on the palms of their hands. They bend their fingers and support the head end of their bodies with their knuckles instead of their open palms.
They are shy, peaceful vegetarians who live in family groups consisting of a dominant adult male with several adult female mates and their children. Subadult males are tolerated in the family as long as they are not actively competing with the dominant male for mates. Gorillas are also very sexually dimorphic
. Adult males average about 350 pounds (160 kg.) and reach 400 pounds (181 kg.) in the wild, while most adult females are only about 155 pounds (70 kg.) and much less muscular.
There are three varieties, or subspecies, of gorillas. The rare mountain gorillas live at high altitude in sparse woodlands in the mountains of Central Africa, while the more numerous lowland gorilla varieties live in the dense forests of West Africa. Most zoo gorillas are of the lowland varieties.
Female lowland gorilla
Gorilla locomotion--video clip by Dennis O'Neil
(length = 1 min.)Gorilla diet--video clip by Dennis O'Neil
(length = 2 mins. 1 sec.)
Common chimpanzee
Peaceful allogrooming
(common chimpanzees)The common chimpanzees more closely resemble humans than do the gorillas. Male chimps grow to 5½ feet (1.7 m.) tall and average about 100 pounds (45 kg.) with 6 foot (1.8 m.) arm spans, while females are usually only around 82 lbs. (37 kg.) and are less muscular. However, like the other large apes, chimpanzees are more comfortable getting around quadrupedally. They are also knuckle walkers like gorillas. The natural habitat of chimpanzees includes both tropical forests and bordering savannas
in Africa.
Chimps are intelligent animals with generally pleasant personalities. However, the males are less peaceful than the smaller females. This behavior difference is typical of most primate species, including humans. The chimpanzee diet is usually at least 90% vegetarian. Males are more likely to eat meat than are females, although chimp hunting skills are relatively poor.
Chimpanzees hunting in trees--video clip from National Geographic Society
(length = 2 mins. 2 secs.)Chimpanzees live in fluid societies of 10-50 individuals. Membership changes through time as females move from one community to another. Males usually stay together and act as a group in defending their mutual territory. However, within each community, males frequently establish short-term alliances in order to compete for dominance. Chimpanzees do not form monogamous mating bonds. Females usually mate with different males throughout their lives.
Bonobos are close relatives of common chimpanzees in the same genus, Pan
. They are sometimes referred to as pygmy chimpanzees. Despite this name, they are only slightly smaller than the common chimps. Bonobos usually have blacker hair with tufts at the side of their faces, longer arms and legs, as well as slimmer bodies. Their vocalizations are also quite different from those of the common chimpanzees. Like many of the Old World monkey species, adult female bonobos normally have prominent "sexual skins." However, unlike monkeys, bonobo females are sexually receptive most of the time and have large sexual skins throughout the year.
Female bonobo Bonobo sexual skin Bonobos have fluid social groupings similar to the common chimpanzees, although bonobos are less excitable and aggressive. Male-female alliances also are more important for bonobos. Older females at times even become group leaders. Bonobos are unique among nonhuman primates in primarily engaging in sexual intercourse face to face. Gorillas do it occasionally. Both heterosexual and homosexual intercourse are common among bonobos. Copulation occurs frequently as a means of reducing tension in the community and has become recreational for them. In this and other traits, bonobos are like humans.
Today, the bonobo range is limited to the forests south of the Zaire River in West Central Africa, and there are considerably fewer of them than the common chimpanzees.
Chimps and Bonobos--an explanation of how they became different species
This link takes you to a video at an external website. To return here, you must click
the "back" button on your browser program. (length = 3 mins, 47 secs)Common chimpanzees are the most successful of all apes in that there are more of them and they have the widest geographic range. However, their numbers have been significantly reduced. A century ago, there were millions of them in the wild. Today, there are less than 200,000. This sharp decline apparently has been mostly due to the rapid increase in human populations and the accompanying natural habitat decimation. An additional factor has been the desire of many people in West and Central Africa to eat chimpanzees and other non-human primate species. At least 4,000 chimpanzees are killed for their meat every year. Chimpanzees and gorillas have suffered devastating Ebola epidemics as well. During 2002 and 2003, approximately 5,000 gorillas succumbed to this highly contagious, almost always fatal disease. The bonobos are at an even greater risk of disappearing since there may be only about 6,000 of them remaining in the wild. Gorillas and both species of chimpanzees have the misfortune of living in nations in which wildlife protection has been severely disrupted by civil wars and the breakdown of effective national authority over the last two decades. It is unlikely that the populations of these apes would be able to spring back rapidly even if they were more carefully protected because they have low reproductive rates. Under the best conditions, adult female chimps usually only have one baby every 5 years. The other great apes are similar.
The great apes are not alone in being threatened by humans. The World Conservation Union estimates that a quarter of all primate species will go extinct within the next 20 years. Many will disappear with little known about them.
This page was last updated on
Friday, February 15, 2008.
Copyright © 1998-2008 by Dennis
O'Neil. All rights reserved.
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